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25th-Century space pilot Ed Mercer divorces his wife Kelly Grayson after he catches her cheating on him. A year later, he accepts a position as Captain of the USS Orville and finds to his dismay that his ex-wife Kelly will serve as his First Officer. During the USS Orville's first mission, the hostile alien Krill Captain attempt to steal a device that can accelerate time, which has both beneficial and dangerous applications. Mercer and Grayson rig the device to destroy itself and the Krill vessel.